Habitat: narrow sandstone box canyon with floodplain, upland woods, hemlock/hardwood coves; moist sandstone
Locality: Conkle's Hollow State Nature Preserve, Big Pine Rd. E of OH 374 between OH 678 and 664 Hocking Hills State Park
Habitat: sandstone bluffs and gorges, creek floodplains; associated with oak, maple and tulip poplar; wet sandstone floor
Locality: Lake Katharine State Nature Preserve W of Jackson
Habitat: upland mixed hardwoods, sandstone bluffs, hemlock/hardwood coves, floodplains, pine forests; soil
Locality: Little Rocky Hollow State Nature Preserve, first hollow W of Kreachbaum Rd. and N of Big Pine Rd.
Habitat: sandstone bluffs and gorges hemlock/hardwood caves, floodplains, upland mixed hardwood forests, pine forests; sandstone stellar cave
Locality: Crane Hollow Nature Preserve, S of Gibsonville N of Conkle's Hollow State Nature Preserve, E of OH 374 W of Cream Ridge Rd.