Habitat: Dry, calcareous upland slopes, Rock Type = Calcareous.; on calcareous boulders.
Locality: Atigun Gorge, Brooks Range, Dalton Hwy. Mountain pass through Brooks Range
Habitat: beech maple forest, along S shore of Acton Lake; limestone boulder, ravine
Locality: Hueston Woods State Park, Hueston Woods (big woods) Interpretive Area
Habitat: narrow sandstone box canyon with floodplain, upland woods, hemlock/hardwood coves; sandstone conglomerate
Locality: Conkle's Hollow State Nature Preserve, Big Pine Rd. E of OH 374 between OH 678 and 664 Hocking Hills state Park