Habitat: sandstone bluffs and gorges hemlock/hardwood caves, floodplains, upland mixed hardwood forests, pine forests; sandstone floor of creek
Locality: Crane Hollow Nature Preserve (crane hollow inc.) S of Gibisonville and N of Conkle's Hollow State Nature Preserve, bordered on E by OH 374 and W by Cream Ridge Rd.
Habitat: mixed mesophytic forest, along Pickle Creek, near campground; dry sandstone rock
Locality: Hawn State Park, ca 60mi S of St. Louis, near Farmington, Hwy 44
Habitat: narrow sandstone box canyon with floodplain, upland woods, hemlock/hardwood coves; base of dogwood
Locality: Conkle's Hollow State Nature Preserve, Big Pine Rd. E of OH 374 between OH 678 and 664 Hocking Hills State Park region