Habitat: Narrow sandstone box canyon, with floodplain, upland Moods, heslock/hardwood coves; associated with staple, hemlock, oak, tulip poplar; Rock Type sandstone, sandstone conglonerate; on sandstone seepage, E. ria.
Locality: U.S.A.: OHIO; Hocking Co., Conkle's Hollow State Nature Preserve, Big Pine Rd., E of Oh 374, between Oh 678 & 664, Hocking Hills State Park region.
Habitat: bottom of ravine; On limestone rocks in stream
Locality: Seven Caves limestone area adjacent to Rocky Fork Cr. Between Hillsboro and Chillecothe, Fern Dell Cave Trail.