Habitat: Sandstone bluffs and gorges ; hemlock /hardwood coves ; floodplains ; upland mixed hardwood forests ; pine forests ; associated with oak ; maple ; tulip poplar hemlock ; pine; rock type = sandstone conglomerats. On exposed clay in roadbed ; with Notothylas ; Anthoceros ; Phaeoceros ; Micromitrium ; Aphanorrhegms.
Locality: Crain Hollow Nature Preserva (Crane Hollow Inc.) ; S of Gibisonville & N of Conkle's Hollow State Naature Preserve ; E of OH 374 & W of Cream Ridge Rd.