Habitat: narrow sandstone box canyon with floodplain, upland woods, hemlock/hardwood coves; grassy soil
Locality: Conkle's Hollow State Nature Preserve, Big Pine Rd. E of OH 374 between OH 678 and 664 Hocking Hills state Park
Habitat: sandstone bluffs and gorges, hemlock/hardwood coves, floodplains, upland nixed hardwood forests, pine forests; soil old field
Locality: Crane Hollow Nature Preserve (Crane Hollow) S of Gibisonville St. N of Conkle's Hollow State Nature Preserve, E of 374 Cream Ridge Road